In his prime he was a “majestic presence” who drew awe from visitors to one of Wales’ top tourist destinations.

But the effects of age catch up with even the king of the jungle with the news that Hugo the 17-year-old lion has died.

Folly Farm Adventure Park and Zoo in Pembrokeshire announced “with heavy hearts” the passing of their beloved male African lion following a short illness.

“Our thoughts are with our keeping team who are feeling his loss most deeply,” it said in a statement.

Hugo arrived at the attraction near Tenby in 2014 with his mate Luna and their four cubs – Sola, Zahra, Alika, and Ebele – and swiftly became a “firm favourite” among staff and visitors.

Following the illness it was discovered he was suffering from chronic kidney disease, a progressive and irreversible condition common in older big cats.

The decision was taken to put him down due to his age, given more episodes of illness were likely.

“It is with very heavy hearts that we announce the passing of our beloved Hugo,” the statement read.

“His recent illness was determined to be an uraemic episode and sadly, more episodes were likely.

“Following an ethical review and given Hugo’s age and the nature of his condition, the difficult but humane decision to provide a dignified goodbye to him was taken to prevent further suffering.”